The article proposed a mechanism called "Predicative Coding" that doesn't require backward flow of information to perform weight adjustment as backward propagation. Futher the author showed that the performance of Predicative Coding is similar to backward propagation.

The author, as such, concluded that predicative coding is a plausible way for biological neuron system to operate.

A nice demo of usage of Ghidra, a reverse engineering tool. The author find and modified game logic to make it possible to marry two people of the same gender.